Topic: Fossil Records

Azjah

Date: 2008-08-29 21:29 EST
Eurypterids

The eurypterids (sea scorpions) include the largest known arthropods that ever lived (with the possible exception of Arthropleuridae). They are members of the extinct class Eurypterida (Arachnomorpha, Chelicerata) and predate the earliest fishes. The largest, such as Jaekelopterus, reached 2 meters or more in length, but most species were less than 20 centimeters. They were formidable predators that thrived in warm shallow water in the Cambrian and survived to the Permian from 510 to 248 million years ago. Although called "sea scorpions", only the earliest ones were marine (most lived in brackish or freshwater), and they were true scorpions. They moved from the sea to fresh water probably by the Pennsylvanian period.

http://i272.photobucket.com/albums/jj171/Azjahh/Geology/Eurypterid.jpg

The typical eurypterid had a large, flat, semicircular carapace, followed by a jointed section, and finally a tapering, flexible tail, most ending with a long spine at the end (Pterygotus, though, had a large flat tail, possibly with a smaller spine). Behind the head of the eurypterids were twelve body segments. These segments are formed by a dorsal plate called tergite, and a ventral plate called sternite. The tail, known as the telson, is spiked in most eurypterids like in modern scorpions and in some species it may have been used to inject venom, but so far there is no certain evidence any eurypterids were venomous. Most eurypterids have paddles toward the end of the carapace and beyond, which were used to propel themselves through water. Some argue that the paddles were also used for digging. It is possible that it was used for both. Underneath, in addition to the pair of swimming appendages the creature had 4 pairs of jointed legs for walking, and two large claws at the front, chelicerae. The walking legs had odd hairs, similar to modern day crabs. Other features, common to ancient and modern arthropods of this type, include one pair of compound eyes and a pair of smaller eyes called ocelli, in between the other larger 2 eyes.

Eurypterid fossils have a near global distribution. Among the largest eurypterids are the Hibbertoptermina, named after the British palaeontolgist S. Hibbert, who described Hibbertopterus scouleri at a limestone quarry in East Kirkton, Scotland, in 1836. Fossil tracks were identified recently in East Lothian, Scotland, as made by a 1.6 meter long Hibbertopterus (Whyte, 2005).

Eurypterids are related to the modern marine horseshoe crabs. About two dozen families of eurypterids are known. They went extinct during the Permian-Triassic extinction 251.4 million years ago. A predatory arthropod whose traces are known as Protichnites, found in Cambrian strata dating from 510 million years ago, is a possible stem group eurypterid, and is among the first evidence of animals on land.

In 2007, a group paleontologists led by Simon Braddy at the University of Bristol discovered a sea scorpion larger than a human being, which was claimed to have been the largest arthropod which ever lived. This discovery was made in a 390 million year old rock containing the fossil of a huge claw or chelicera.

Azjah

Date: 2008-08-29 21:35 EST
Pelycosaur http://www.ucmp.berkeley.edu/synapsids/pelycosaurs .html

The "pelycosaurs" are members of the Synapsida, a major branch of the Amniota, or egg-laying tetrapods. Pelycosaurs are the earliest, most primitive synapsids, a group characterized by a single dermal opening in the skull permitting muscle attachment to the jaw. This means that the pelycosaurs are not reptiles, since reptiles have two such openings in their skulls. Like most other groups of synapsids, all pelycosaurs are now extinct. In fact, the only currently living synapsids are the mammals. It is believed that the pelycosaurs, like their living mammal relatives, were endothermic, which means that they maintained a constant internal body temperature. This is another characteristic that sets pelycosaurs apart from the reptiles. If this view is correct, then pelycosaurs are one of the earliest examples of endothermic animals.

In many respects, the pelycosaurs are intermediate between the reptiles and mammals, and so they have commonly been referred to as "mammal-like reptiles". The pelycosaurs indeed resemble large lizards in their overall appearance, but as we have seen, this is a misnomer since pelycosaurs are not reptiles. The name "pelycosaur" refers to a paraphyletic group of basal synapsids consisting of six separate branches: Caseasauria, Varanopseidae, Ophiacodontidae, Edaphosauridae, "Haptodonts", and Sphenacodontidae. It is currently believed that haptodonts themselves are a paraphyletic group. The Sphenacodontidae are probably the most closely related to the Therapsida (the group that includes mammals).

Pelycosaurs first appeared during the upper Carboniferous (Lower Pennsylvanian) and went extinct by the end of the Permian. With the formation of the super-continent Pangea in the Permian, continental area exceeded that of oceanic area for the first time in geological history. The result of this new global configuration was the extensive development and diversification of Permian terrestrial vertebrate fauna including pelycosaurs. Consequently, their fossils are found in multiple localities around the world. However, the majority of pelycosaur fossils have been discovered in Texas and Oklahoma, where they are among the most common fossils of the Early Permian. Much of the oil in the two states derives from the Permian period, and the search for this oil is partially responsible for the plethora of pelycosaur fossils found.

http://i272.photobucket.com/albums/jj171/Azjahh/Geology/Pelycosaur1.jpg

Dimetrodon is the most familiar example of a pelycosaur. A powerful carnivore, Dimetrodon was the top predator of the Permian period. With its large head and numerous large, sharp teeth, it fed on other vertebrates, including other pelycosaurs that shared the swamps where it lived. Dimetrodon was about three-and-a-half meters (eleven feet) long and probably weighed about 100-150 kg (200-300 pounds). Four short and thick legs supported this massive body.

The physical appearance of Dimetrodon is quite distinct. While its body was long, it was relatively short. It was built like a large lizard with one important exception: a "sail". The bony sail supports rose nearly a meter vertically from its back and were connected by highly vascularized skin. The most popular conjecture regarding the function of the sail is thermoregulation. Such a sail would allow a large area for acquiring or releasing heat. Studies have shown that the area of the sail is proportional to the volume of the Dimetrodon body, as would be expected if the sail were important for heat exchange. While many tetrapods in many lineages have evolved similar sail-like structures, Dimetrodon is the best known so the majority of work done exploring the function of the sail has centered on this pelycosaur.

However, It must be noted that not all pelycosaurs had sails. Still, most pelycosaurs were similar to Dimetrodon in that they also were carnivores with large, powerful jaws, and two types of teeth: sharp canines and shearing teeth. Two major exceptions to this meat diet were the Caseidae and the Edaphosauridae, who were herbivores. While the Caseids were far more diverse, the most familiar of the herbivorous pelycosaurs is another sail-back, Edaphosaurus. Edaphosaurus had a smaller skull than Dimetrodon, and had large peg-like teeth for grinding and crushing plants. This adaptation allowed pelycosaurs to flourish, and by the beginning of the Permian, pelycosaur genera account for 70 percent of all the known amniotes, outnumbering the reptiles.

Pelycosaurs are an important lineage preserved in the fossil record. They are the earliest known synapsids, the first to evolve specializations that would play an important role in the rise to mammals. They are thus a transitional group in both form and functions. Schad (1994) presents two general rules concerning transitional areas. Firstly, no intermediate form represents a totally intermediate character. This is to say that no evolutionary condition has been found in any taxon where all characteristics are present of both taxa that the condition is intermediate to. Secondly, evolutionary transformation predominates in the extremities, and only later affects the axial system. Bearing in mind these principles, we can begin to understand the development of mammals from their pelycosaur roots.

The pelycosaurs exhibit the first substantial progress of crawling to running. This evolution in the extremities required a modification of the metabolism in the muscular system to provide the energy reuired for more strenuous activity. The resulting change in the axial system brought about endothermy. Supporting this idea is the fact that as later pelycosaurs and later synapsids evolved, the surface area of sail to body mass ratio decreased. This shows the trend of reduced need for outside thermoregulation, which would require an increased use of endothermy, an important characteristic today separating the reptiles and mammals.

http://i272.photobucket.com/albums/jj171/Azjahh/Geology/pelycosaur2.jpg

Azjah

Date: 2008-08-30 16:12 EST
Pyllum Porifera ? The Sponges
Precambrian to present

Fossil sponges have existed since the Precambrian era. However earlier sponges were more likely to be vase or cone shaped whereas today?s sponges are more likely to be massive or branching. Sponges lack true tissues. They have no nerves, muscles or glands. They are composed of needle-like, mineralized spicules and sponging, a fibrous horny substance. In most sponges, the spicules are not joined, and when the sponge dies, the spicules disperse. The inly sponges likely to be preserved are these whose spicules are fused unto a rigid framework.

Simplest sponges are vase-shaped, with pores in the walls through which water passes to the interior. Chambers inside are lined with collared cells bearing small flagella. They beat the water and small bits of food are captured by the collars. There are only two layers of cells in a sponge with a layer of jelly-like material between them that holds the skeleton.

Sponges have no means of movement, there for they generally anchor to a firm object. They vary in size from less than 25mm ? 61cm across. (<1 inch - >2 feet).

Class Demospongia: are what most living sponges are. The skeleton consists of sponging alone or of sponging and siliceous spicules. The spicules are sometimes straight rods, but may have 4 rays diverging at angles of 60 degrees or 120 degrees. Walls of these sponges are commonly thick and folded.

Class Hexactinellida: are glass sponges that have skeletons consisting only of siliceous spicules that have 6 or fewer rays that diverge at 90 degree angles. These spicules are often fused into a rigid framework. This class is sometimes called Hyalospongea.

Class Calcarea: These are the most primitive sponges, with skeletons consisting only of calcareous spicules, usually fused into a rigid framework or of beadlike spheres or flakes. This class is sometimes called Calcispongia.

Class Stromatoporoidea: These were previously class as cnidarians, but the change to sponges is due to their striking similarity with modern sclerosponges. These were important reef builders from the Silurian and Devonian. Stromatoporoids are massive, encrusting, or cylindrical. Most have a layered structure. The skeletons consist of separate or joined horizontal and vertical elements. The average diameter of skeletal masses is only about 4? (102mm) but they may grow to several meters wide. Identifying Stromatoporoids to genus requires specialized techniques.

Precambrian

Cambrian

Protospongia (Hexactinellida)
Lower and middle Cambrian
Widespread in North America.
This is one of the earliest sponges in the fossil record, and one of the simplest. It was an ascon, a single vaselike form, hollow in the middle with an opening at the top. Grew to 19mm long and wide, the spicules are 6mm long, usually found separated, simple, and may have formed reticulated meshwork.

Actinostroma (Stromatoporoids)
Cambrian ? Mississippian
Distribution: Widespread
Description: 2 1/8? (54mm) long, 1 ?? (32mm) wide. Massive form, with dense skeletal deposits. The surface may show small, shallow grooves radiating from tiny pores.

Ordovician

Astylospongia (Demospongia)
Ordovician ? Silurian
Distribution of fossils: New York, Ontario, Indiana, Kentucky, Tennessee, Illinois, Iowa. Two inches long and wide (51 mm)
Especial interest are the vertical and irregular grooves where most likely canals through which the water traveled and was covered over by tissue when the sponge was alive.

Dystactospongia (Demospongia)
Ordovician ? Silurian
Distribution: Virginia, Ohio, Tennessee, Illinois
Tiny radiating canals are visible if the surface is eroded. Very narrow distribution, only commonly found in local abundance rather than widely dispersed.

Hindia (Demospongia)
Ordovician through Permian
Distribution of fossils: Eastern Canada, New York, New Jersey, Maryland, Virginia, Georgia, Ohio, Kentucky, Tennessee, Wisconsin, Illinois, Missouri, Minnesota, Iowa, Nevada.
Description: Fossils have spherical, smooth surfaces with very tiny pores. Stalk and depressions are absent in this sponge. This may indicate that Hindia may have been adapted to roll about freely on the sea floor.

Brachiospongia (Hexactinellida)
Ordovician
Distribution: Ontario, Ohio, Kentucky, Scotland (no where else in the world)
Description: This sponge was 105mm long by 95mm wide with radiating projections. The periphery was divided into 8-12 fingerlike lobes that curved downward, probably forming a base. The upper surface has fairly large, round, elevated openings connecting to hollow loves. Walls were thick, and no root tuft has been found. The form of this sponge is unusual with the radiating lobes.

Aulacera (Stromatoporoidea)
Ordovician
Distribution: Widespread
Description: 4 7/8? (124mm) long, 1 3/8? (35mm) wide. Solid, cylindrical mass, surface is covered with wrinkles or bumps. The cross section shows a large vertical canal, divided horizontally by strongly arched, sheetlike platforms and surrounded by irregular layers of cystlike plates.

Silurian

Astraeospongium (Calcarea)
Silurian ? Devonian
Distribution: New York, Ohio, Kentucky, Tennessee, Illinois, Iowa
Description: 2.5? long and wide (64mm) Saucer shaped with no trace of attachments. Skeleton is felt like mass of 8 rayed spicules, 6 rays in one plane, and 2 perpendicular. The flowerlike spicules are easily visible and make this a distinctive genus.

Stromatopora (Stromatoporoidea)
Silurian ? Devonian
Distribution: Widespread
Description: 3? (76mm) long, 3.5? (89mm) wide as massive or sheetlike calcareous skeletal deposits, sometimes hemispherical. Surface usually has bumps which may be low or high, and may have prominent pits in the centers. Stromatopora means ?layer-pores? and refers to the layered structure.

Parallelopora (Stromatoporoidea)
Silurian ? Devonian
Distribution: Widespread
Description: 2? (51mm) long, 1 ?? (32mm) wide. Massive, sheetlike, or encrusting calcareous skeletal deposit with irregular surface, sometimes covered with distinct bumps. Some species covered with small, shallow grooves radiating from tiny pits. Horizontal layers are prominent and closely spaced.

Devonian

Cliona (Demospongia)
Devonian ? Present
Fossilizes only as borings in brachiopod shells. Today, they are bright yellow. The borings of Cliona help break down shells

Hydnoceras (Hexactinellida)
Upper Devonian ? Pennsylvanian
Distribution New York and Pennsylvania.
Description: This glass sponge is very distantly related to living glass sponges. The siliceous spicules form a strong, tubular skeleton about 21 cm long and 9 cm wide. They were large, and roughly conical with 8 pronounced, sharp, vertical ridges crossed by horizontal ridges.

Prismodictya (Hexactinellida)
Upper Devonian ? Mississippian
Distribution: New York, Indiana
Description: 95mm long, 41mm wide (3.5? x 1 5/8?) as a tall, narrow, vaselike structure with 8 prismlike sides.

Mississippian

Pennsylvanian

Girtyocoelia (Calcarea)
Pennsylvanian
Distribution: Kansas, Oklahoma, Texas
Description: This is a genus distinct from Girtycoelia, which is also Pennsylvanian and found in Kansas and Texas. Girtyocoelia is 70mm long, 19 mm wide. Small, with linear series of distinct hollow spheres pierced by large axial tube. The outer wall is covered with few large, spoutlike pores. In older books, Girtyocoelia is called Heterocoelia.

Permian

Triassic

Jurassic

Cretaceous

Paleocene

Eocene

Oligocene

Miocene